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1、JVIAcceptedManuscriptPostedOnline16September2015J.Virol.doi:10.1128/JVI.01755-15Copyright?2015,AmericanSocietyforMicrobiology.AllRightsReserved.1Evidenceforanancestralassociationofhumancoronavirus229Ewithbats21,2#3,4#1,5#3VictorMaxCorman,HeatherJ.Baldwin,Adriana

2、FumieTateno,RodrigoMelim1,56664Zerbinati,AugustinaAnnan,MichaelOwusu,EvansEwaldNkrumah,GaelDarren7883,95Maganga,SamuelOppong,YawAdu-Sarkodie,PeterVallo,LuizVicenteRibeiro5,107,1112,13146FerreiradaSilvaFilho,EricM.Leroy,VolkerThiel,LiavanderHoek,LeoL.M.153,161,2*

3、1,2*7Poon,MarcoTschapka,ChristianDrosten,JanFelixDrexler891:InstituteofVirology,UniversityofBonnMedicalCentre,Bonn,Germany102:GermanCentreforInfectionResearch(DZIF),partnersiteBonn-Cologne,Germany113:InstituteofEvolutionaryEcologyandConservationGenomics,Universi

4、tyofUlm,Ulm,12Germany134:DepartmentofBiologicalSciences,MacquarieUniversity,Sydney,NSW2109,Australia145:LaboratóriodeVirologia(LIMHC-FMUSP),InstitutodeMedicinaTropicaldeS?oPaulo,15UniversidadedeS?oPaulo166:KumasiCentreforCollaborativeResearchinTropicalMedicine(K

5、CCR),Kumasi,Ghana177:CentreInternationaldeRecherchesMédicalesdeFranceville,Franceville,Gabon188:KwameNkrumahUniversityofScienceandTechnology,Kumasi,Ghana199:InstituteofVertebrateBiology,AcademyofSciencesoftheCzechRepublic,v.v.i.,Brno2010:InstitutodaCrian?a,Hospi

6、taldasClínicasdaFMUSP,S?oPaulo,Brazil2111:InstitutdeRecherchepourleDéveloppement,UMR224(MIVEGEC),IRD/CNRS/UM1,22Montpellier,France2312:DepartmentofInfectiousDiseasesandPathobiology,VetsuisseFacultyBern,University24ofBern,Switzerland2513:FederalDepartmentofHomeAf

7、fairs,InstituteofVirologyandImmunology,Bernand26Mittelh?usern,Switzerland12714:DepartmentofMedicalMicrobiology,AcademicMedicalCenteroftheUniversityof28Amsterdam,Amsterdam,TheNetherlands2915:SchoolofPublicHealth,TheUniversityofHongKong,HongKong,Special30Administr

8、ativeRegion,People’sRepublicofChina3116:SmithsonianTropicalResearchInstitute,Balboa,Panama3233#Theseauthorscontributedequally34*Addressesforcorrespondence:35Professor